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Draw rune symbols with the mouse to build towers (just hold the left mouse button and start dragging)
The runes are shown on the panel at the bottom. You can upgade and sell using the same system.
Collect the souls (your money) by mousing over them once the creeps die. More Information...
You use the mouse to draw symbols that will summon/upgrade/sell towers! Not only that, you get to collect those little creepy guys souls after your towers blast them!